A Flyway Through the Heart of Asia: ‘Heuglin’s Gulls’ Reveal a Hidden Migratory Passage
A GPS-tracked Lesser Black‑backed Gull (Heuglin’s Gull) that helped uncover a transcontinental migration passage linking the Indian Ocean with the Arctic—one of the most significant seabird flyway findings in the Central Asian Flyway to date. Image courtesy of the CAF–Sri Lanka Waterbird Tracking Project

A Flyway Through the Heart of Asia: ‘Heuglin’s Gulls’ Reveal a Hidden Migratory Passage

New GPS tracking uncovers a transcontinental seabird route connecting the Indian and Arctic Oceans – offering a rare glimpse into one of Asia’s most elusive avian journeys.


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A tropical gull with Arctic ambitions

When we think of long-distance bird migration, our minds often conjure images of sandpipers flying nonstop across oceans or raptors riding thermals between continents. But one of the most overlooked players in this global phenomenon is the Lesser Black‑backed Gull (Heuglin’s Gull) (Larus fuscus heuglini), a large-bodied coastal gull that winters in the tropics and breeds in the high Arctic. Now, a new study has shed light on its remarkable journey – revealing an unexpected migratory corridor that bisects the vast Eurasian landmass.

Close-up of a tracked Lesser Black‑backed Gull (Heuglin’s Gull) prior to release. Tagging efforts focused on adult individuals wintering in tropical South Asia. Image courtesy of the CAF–Sri Lanka Waterbird Tracking Project

Using GPS-GSM tags, researchers from Sri Lanka, China, and India tracked the movements of three Lesser Black‑backed Gulls (Heuglin’s Gull) between 2021 and 2023. What they discovered was not a direct flight along the shortest route, nor a maritime cruise hugging coastlines – but an intermediate path that blends coastal and overland strategies, traversing thousands of kilometres from the Indian Ocean to the Russian Arctic.

A tagged Lesser Black‑backed Gull (Heuglin’s Gull) moments after release in Sri Lanka. This individual later completed a full migration to the Russian Arctic, helping define a new migratory corridor across Eurasia. Image courtesy of the CAF–Sri Lanka Waterbird Tracking Project

A newly charted route through the Central Asian Flyway

Tagged on Sri Lanka’s Mannar Island, the gulls first crossed to India’s western coast before navigating northwards through Pakistan, Afghanistan, and Central Asia – eventually reaching the Yamal Peninsula in Arctic Russia. Their route, totalling around 8,000 km, bypassed the theoretically shorter trans-Himalayan corridor (6,346 km) and the far longer maritime route via the Suez and Atlantic (15,000 km). This deviation increased the overall migration distance by approximately 30%, but spared the birds the cost of flying over extreme elevations such as the Himalayas (over 6,000 m a.s.l.).

Migration routes of Lesser Black‑backed Gulls (Heuglin’s Gull) tracked from Sri Lanka to Arctic Russia between 2021 and 2023. Each coloured line represents a different individual and season, revealing consistent use of a transcontinental passage through inland South and Central Asia. Circles mark stopover locations, with size proportional to duration. Breeding and non-breeding sites are indicated by triangles. Image courtesy of CAF–Sri Lanka Waterbird Tracking Project

Instead, they flew mostly over lowlands, averaging just 323 m above sea level. Only 9% of their flights crossed ground elevations above 1,000 m. By following river systems and wetland-rich landscapes – particularly the Ob River basin – the birds likely found suitable stopover sites and avoided the physiological demands of high-altitude flight.

A migration fuelled by flexibility

The gulls exhibited a flexible mix of travel strategies. Northbound, they made more stopovers in Central Asia. Southbound, they bypassed these regions entirely, suggesting either better fuelling from Arctic breeding grounds or favourable tailwinds. Interestingly, their stopover duration – not actual flight speed – had the strongest influence on total migration time.

While the birds mostly flew during the day (88% of all movement), night flights played a surprising role during southward migration, comprising up to a third of the total migration distance. This contrasts with other large gulls and may hint at an adaptive response to harsh or resource-scarce regions.

Lesser Black‑backed Gulls (Heuglin’s Gull) also appear to use a “fly-and-forage” strategy – soaring when possible and searching for feeding opportunities en route. This likely explains why their average instantaneous speed was relatively high (~47 km/h), but their overall daily progress more modest (409 km/day). The balance of speed, rest, and foraging seems finely tuned to their ecological context.

Implications for conservation and research

This study is the first to document a consistent transcontinental migration route for a seabird in the Central Asian Flyway, long considered one of the least understood of the world’s major migratory corridors. By uncovering this passage, the research not only advances our understanding of gull migration ecology but highlights the broader significance of Eurasian wetlands, coasts, and inland stopovers for long-distance migrants.

The findings carry urgent conservation implications. Tropical coastal zones – such as those used by wintering Lesser Black‑backed Gulls (Heuglin’s Gull) – are under mounting pressure from coastal development, pollution, and climate change. Protecting these areas is essential not just for gulls but for dozens of other migratory species that rely on similar corridors.

Moreover, the study contributes to growing evidence that some seabirds, traditionally thought to favour oceanic routes, can and do adapt to overland migration when topography and resources allow. The observed overlap with the Steppe Gull (Larus fuscus barabensis) – a closely related subspecies – raises new questions about evolutionary divergence and the shaping of migratory strategies.

A window into unseen journeys

As the authors note, the Lesser Black‑backed Gull (Heuglin’s Gull) remains one of the least understood Arctic-breeding gulls. Thanks to advances in GPS tracking and international collaboration, this gap is slowly closing. The study underscores the power of modern technology to reveal not only where birds go, but how they make decisions in the face of geography, energy demands, and environmental change.

Professor Sampath Seneviratne, the study’s senior author, hinted that further insights are on the way. A companion paper on Brown-headed Gulls crossing the Trans-Himalaya is currently in press. Together, these studies mark a turning point in our understanding of how birds bridge the ecological extremes of the tropics and the Arctic.


References

Panagoda, G., Wijethunge, I. K., Zhang, B., Meng, F., Liu, Y., Kotagama, S., Mundkur, T., Balachandran, S., Cao, L., & Seneviratne, S. S. (2025). A transcontinental migratory passage linking the Indian Ocean with the Arctic Ocean: Migration of Heuglin’s Gulls from tropics to Arctic. Biotropica, 57:e70045. https://doi.org/10.1111/btp.70045

AviList Core Team. (2025). AviList: The Global Avian Checklist, Version 2025. https://doi.org/10.2173/avilist.v2025


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